Revision of the Shield-bug Genera Holcostethus Fieber and Peribalus Mulsant et Rey (Heteroptera, Pentatomidae) of the Palaearctic Region

摘要

A complex of the heteropteran genera centering around Peribalus Mulsant et Rey and Holcostethus Fieber is considered. The genus Dryadocoris Kirkaldy reveals no relationship with the above genera and is believed to represent a separate clade of the family Pentatomidae. The genera Peribalus and Holcostethus are revised. The former includes three subgenera: Peribalus s. str. with two species, Asioperibalus subgen. n. (type species Ci-mex inclusus Dohrn) with six species, and Tianocoris subgen. n. (typespecies Holcostethus manifestos Kiri-tshenko) with two species. Holcostethus embraces two subgenera: Holcostethus s. str. and the monotypic Enigmo-coris subgen. n. (type species H. fissiceps Horvath). Two new species are described: Peribalus tianshanicussp. n. from the Tien Shan Mts. and P. przewalskii sp. n. from the northern part of China (Huan He River). P. capitatus Jakovlev and P. vernalis (Wolff) are downgraded to subspecies of P. strictus (F.). P. ovatus Jakovlev is syn-onymized with P. inclusus(Dohrn). Two new monotypic genera related to the revised complex of genera are established, Paraholcostethus gen. n. (type species Peribalus breviceps Horvath) and Himalayastethus gen. n. (type species H. pilosus sp. n. from Kashmir). A key to, and morphometric characters for all the taxa considered are provided. The key characters, including both male and female genitalia, are illustrated, and distributional maps are given.