Applicator for applying a product on keratinous fibers, in particular eyelashes or eyebrows, comprising a support (20; 30; 40; 91; 101; 111; 120; 130; 140; 151; 180, 181, 182 ; 190) and bristles secured to said support for product application, wherein the support is not twisted and has at least one through hole (21) of non-constant cross section in which is held a tuft of hairs (22 ).
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