[1]National Parks Wildlife and Plants Conservation Department Bangkok 10900 Thailand;
[2]The UniversityMuseum The University of Tokyo Bunkyo Tokyo 113-8654 Japan;
[3]Graduate School of Agricultural and Life Sciences The University of Tokyo Bunkyo Tokyo 113-8654 Japan;
[4]Yamashina Institute for Ornithology Abiko Chiba 270-1145 Japan;