Laboratory of Marine Organism Taxonomy and Phylogeny Shandong ProvinceKey Laboratory of Experimental Marine Biology Institute of Oceanology Center for Ocean Mega-Science ChineseAcademy of Sciences Qingdao 266071 ChinaLaboratory for Marine Biology and Biotechnology Pilot NationalLaboratory for Marine Science and Technology(Qingdao) Qingdao 266237 ChinaUniversity of Chinese Academy of Sciences Beijing 100049 China;