Hangzhou Key Laboratory for Ecosystem Protection and Restoration College of Life and Environmental Sciences Hangzhou Normal University Hangzhou 311121 China;
Jiangsu Key Laboratory for Biodiversity and Biotechnology College of Life Sciences Nanjing Normal University Nanjing 210023 China;
Hainan Key Laboratory for Herpetological Research College of Fisheries and Life Science Hainan Tropical Ocean University Sanya 572022 China;