Laboratory of Marine Organism Taxonomy and Phylogeny Shandong Province Key Laboratory of Experimental Marine Biology Institute of Oceanology Center for Ocean Mega-Science Chinese Academy of Sciences Qingdao 266071 China;
Laboratory for Marine Biology and Biotechnology Pilot National Laboratory for Marine Science and Technology(Qingdao) Qingdao 266237 China;
University of Chinese Academy of Sciences Beijing 100049 China;