Nature ConservLition of Snake Island and Laotleshan Mountain 116041 Liaoning;
Chengdu Institute of Biology Academy of Sciences 610041;
College of Life Sciences and Food Engineering University 644007;
Key Laboratory of Southwest China Wildlife Resources Conservation(Ministry of Education) West Normal University 637009;
group living; inbreeding risk; pairwise relatedness; snakes; Gloydius shedaoensis;